This is a long shot and you may have done it already, but

In the home folder /Library/ preferences are two mail preferences. It might be worth taking them out and restarting the computer. If it doesn’t help, you can put them back into the folder
com.apple.mail
com.apple.mail-shared
I will let the powers that be on Fine Tuned Mac comment if it is advisable. As I said you may have already tried it